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Four girls have different reasons for joining the Women's Land Army: patriotic Joyce wants to 'do her bit', argumentative Nancy is there under sufferance, and sensible Annie joined so that her younger sister Bea could escape an abusive father. Bea rebels from her sister's control and finds herself enthralled by a charming GI, Cal Gillespie, while at the dance Nancy tries to get closer to Lord Hoxley.
Bea has been hiding her pregnancy from everyone for three months, but it is becoming increasingly difficult. Tucker is on the prowl for enemy sympathisers and Nancy is in his sights. Farmer Finch gets involved in a scam to flog home-made carrot whiskey, while Lord Hoxley realises that his marriage is over - and this drives him into the arms of Nancy. Meanwhile, Tucker finds Annie to tell her the devastating news that her husband has been killed in action.
Three months on, and Annie is not dealing very well with Harry's death. Nancy and Lawrence's attraction has blossomed into a full-on affair. Meanwhile, Esther is screening Nancy's letters for Sergeant Tucker, fearing that she might be a Nazi collaborator. Joyce, the patriot, has a dilemma - should she shelter her deserter husband or turn him in? Farmer Finch's old enemy Ernest Luckhurst, a food rations officer, arrives at the farm and is intent on catching him out.
Farmer Finch hires some 'trekkers', families displaced by the bombing, as cheap labour, but when there is then a break-in at the manor house Lady Hoxley orders Finch to get rid of them all. A riot ensues and Tucker gets the dogs out. The Hoxleys hold a swanky party, but after clashing with Nancy over a visit to her husband John, Joyce tells Lady Hoxley about the affair. Billy finally proposes to Bea, but she goes into labour before she can answer.
Bea prepares for her wedding to Billy and discovers that Finch has doubts about her motives. Determined to leave, Nancy is shocked to the core when Ellen reveals Lawrence's dark secret. Will Annie choose her duty to Harry or her love for Adam? Meanwhile, Tucker sets out on a trail of murderous revenge that will have shocking consequences for Nancy, Annie, Bea and Joyce.